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Who put the frogs backward on the G5 pony models?

Non-Fungible Trixie -
Magical Inkwell - Wrote MLP fanfiction consisting of at least around 1.5k words, and has a verified link to the platform of their choice

As an example, note how the point of the heart-frog on Zipp’s hoof points backward in this gif:
When screwed on properly, the bulbs/lobes of the heard should instead be located toward the posterior of the hoof while the point of the heart/arrow should point to the front of the hoof.

Note how Izzy is presumably rearing. The hearts that form her frogs should point upward.
What puzzles me is that this isn’t some mistake that can be chalked up to no one in the animation studio having ever seen an IRL horse before. The frog is a part of horse anatomy I would not expect them to add unless they already knew horses were supposed to have them. Stylizing them into hearts makes perfect sense in the visual language of Equestria. However, somehow their hearts pointing the opposite direction of IRL horses got the greenlight for their generic pony model early in the process because the decision was remarkably consistent.
It would be one thing if this were just some artistic decision, like stylizing their heads. However, I fear a number of new artists will copy G5’s artistic direction, mistakes and all, and be hoodwinked into drawing backward frogs on their ponies without ever knowing better and citing G5 official media when the discrepancy is explained to them.
Just Wayne
Elements of Harmony - Had an OC in the 2022 Community Collab

Umbreon is best Pokemon!
Obviously it’s done to have some sort of kid appeal. Putting cute little hearts on ponies to let you know they’re indeed cute little ponies full of love. It’s like Care Bears really but at least more clever and subtle. They wouldn’t want to put upside down heart on their ponies because that would symbolize the opposite of love after all. :P
It’s especially obvious with this scene. Heart hoof means they can put a heart symbol at the end of the movie.
And to be fair, the original G4 ponies didn’t get their frogs until the movie anyway. They’re all just little stubby cylinders that are completely flat at the bottom. :P
Background Pony #653B
If the shadowy alicorn has heart-shaped hooves, I’ll love her even more.
Just Wayne
Elements of Harmony - Had an OC in the 2022 Community Collab

Umbreon is best Pokemon!
Oh and I just realized there is this “Hooves to heart” thing they say, so I guess that makes sense too. :P
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